Corresponding author: Steven L. Wick ( utpaleontology@gmail.com ) Academic editor: Oliver Rauhut
© 2021 Steven L. Wick.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Wick SL (2021) Paleontological inventory of Paleozoic, Late Mesozoic, and Cenozoic plant, invertebrate, and vertebrate fossil species from Big Bend National Park, Texas, USA - over a century of paleontological discovery. Zitteliana 95: 95-134. https://doi.org/10.3897/zitteliana.95.73026
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The extraordinary paleontological record from Big Bend National Park (BIBE), Texas chronicles nearly 120 million years of largely uninterrupted deposition through Late Cretaceous, Paleogene and Neogene time. Therefore, the park records one of the most complete and continuous fossil records of its kind in North America, if not the world. Paleontologists have collected and studied fossils from BIBE for over a century and nearly 1400 fossil species have been reported thus far. The BIBE paleontological record includes type specimens representing 44 scientifically valid species (five plants, nine invertebrates, and 30 vertebrates). Numerous other reported specimens are very likely new to science but have yet to be formally named. The present catalog presents the currently known assemblage of fossil plant, invertebrate, and vertebrate species from BIBE within a single, comprehensive record with significant references for each. This work is designed and written to be a research and resource management tool for scientists and non-scientists alike.
Cretaceous, Neogene, paleobiodiversity, Paleogene, paleontology, taxonomy
For more than 100 years, paleontological researchers have made some of North America's most important fossil discoveries in the Big Bend region of West Texas, USA - many of those in what is now Big Bend National Park (BIBE) (Fig.
The updated taxonomic catalog herein is derived from a seminal paleontological inventory of Big Bend National Park produced by
Generalized stratigraphic column (A) exposed within Big Bend National Park, Texas, USA (B). Approximate absolute stratigraphic ages are based upon biostratigraphic and radiometric information from multiple sources (
The original 2015 (unpublished) catalog was developed as an internal NPS document so that NPS interpretive and law enforcement personnel, resource managers, and qualified permitted academics might better explain, protect, manage, and research the diversity and significance of the park's fossil resources. Hence, it was written using uncomplicated language so that it could be better understood by readers with variable levels of interest and expertise. That approach is maintained here. Whatever the case, it must be noted that this catalog (like all projects of its type) remains a work-in-progress. New discoveries will undoubtedly add to the park's paleobiodiversity and new explorers will emerge over the coming decades to expand upon what we have discovered thus far. It must also be noted that several fossil species relevant to the BIBE paleontological story have been discovered just outside of the park in the same geologic formations exposed within it. These were also included in the present catalog under the assumption that these species are very likely present in the park as well but have yet to be found there.
Relevant references involving the various individual species reported here is provided within each of the taxonomic lists and so specific references are not included within the preliminary text. Repositories and accession numbers for the specimens representing the species listed in the catalog can be found in their respective referenced works. Furthermore, understanding the changing landscape of Big Bend is critical to understanding its paleontological story. The reader is, therefore, strongly encouraged to review
Fossils from Big Bend National Park are widespread within Mesozoic and Cenozoic strata which are well-exposed throughout the park. Paleozoic strata are not well exposed within the park and are largely confined to its northern margins and so fossils from this time are not well known. These older rocks were laid down some 330-285 million years ago then subsequently deformed during the Ouachita orogeny. They appear in the configuration that we see today as the subsequent result of Laramide compression, faulting, and erosion during more recent times (e.g.,
Around 120 million years ago, a warm, shallow sea (the Western Interior Seaway) bisected North America dividing the continent in half from today's Gulf of Mexico to the Arctic Ocean (
Approximately 90 million years ago, the shallow Cretaceous seaway began a gradual retreat to its present location - today's Gulf of Mexico. Calcareous marine muds, and silty clay containing more terrigenously-derived sediments were deposited on the nearby shallow, marine shelf along with the remains of giant bivalves, oysters, sharks, fish, ammonites, and mosasaurs. Gulfian Series limestones and shales of the flaggy Boquillas Formation and soft bentonitic clays of the Pen Formation were deposited during this time (
Around 78 million years ago, Big Bend was situated upon the shore of the ancient seaway (
Some 70 to 65 million years ago, Laramide tectonism began uplifting the proto-Rocky Mountains to the west. As a result, the Late Cretaceous shoreline had retreated well to the east of today's park (
The end of the Cretaceous Period was also a time of great change for life on Earth. Although there are several hypotheses for the extinction of the dinosaurs some 66 million years ago, their disappearance at the end of the Cretaceous gave rise to the 'Age' of mammals. Whether caused by climate change, disease, or the impact of a large meteor in the Yucatan of Mexico, this extinction event occurred during deposition of the Black Peaks formation in BIBE, one of the few public lands in North America which contain strata that span the Cretaceous-Paleogene (K-Pg) extinction boundary.
Around 63 million years ago (Paleocene time) the dinosaurs were gone. However, ancient mammals survived the K-Pg extinction event (as did avian dinosaurs - the birds) and began to evolve on the ancient river floodplains in BIBE. Although this was the same river system which originated millions of years earlier during Javelina Formation time, the Rocky Mountains continued their unrelenting uplift (e.g.,
55 million years ago during early Eocene time, the Tornillo Basin continued to aggrade with fluvial sediments of the Hannold Hill Formation in BIBE (
During mid-Eocene time (about 46 million years ago), the Laramide Orogeny had almost reached its culmination and the Big Bend region was now elevated several thousand feet. Erosion then became the dominant regime, stripping away much of the Hannold Hill, Black Peaks, Javelina, and Aguja formations throughout BIBE and surrounds. All of these strata (as well as the fossils preserved within) might have been lost. However, what remained of them was then covered by deposits laid down by a new river system that developed atop the ancient, infilled basin. As a result, sediments of the Canoe Formation were laid down unconformably on the previously eroded surface. These new rocks were made up of thick fluvial channel sands and gravels (the Big Yellow Sandstone in the park) as part of a braided river system (
Approximately 42 million to 32 million years ago (during middle Eocene to early Oligocene time) Big Bend experienced a strikingly different depositional regime as widespread volcanism commenced. Strata deposited during this time differ markedly across the region as the result of the changing loci and composition of various igneous intrusions, lavas, ash-falls, as well as the fluvial volcaniclastics derived from them via weathering (
During early Oligocene time (around 32 to 26 million years ago), volcanism continued with a series of eruptions in what is today BIBE (
By the end of Oligocene time (around 20 million years ago), the Rocky Mountains stood in bold relief above the western plains. Compressional stresses involved in mountain-building finally eased across the North American continent resulting in a 'relaxation' of continental crust. As a result of this trans-continental stretching, rift zones developed which, over time, allowed large bodies of rock to slide downward along active faults, producing a horst-and-graben topography. This created the North American, Basin and Range Province which spans southern Canada to northern Mexico including Big Bend. The Big Bend region saw the development of several grabens and resultant bolsons including one within the central part of today's BIBE (from the Sierra del Carmen to the east to the Mesa de Anguila to the west). This graben formed a "sunken block" of strata, down-dropped several thousand feet by faulting (
Eventually, similar bolsons throughout west Texas were infilled and subsequently linked by the Rio Grande (achieving through-flow to the Gulf of Mexico only within the last 2 million years or so). Once established, the downcutting Rio Grande and its tributaries (forerunners of today's Terlingua and Tornillo creeks within BIBE) largely gutted the infilled bolsons during Pleistocene time leaving only remnants of them today. The Rio Grande is the youngest major river system in the United States and continues to serve as the principal erosional conduit in the region.
The geologic formations within BIBE vary widely regarding composition, thickness, depositional environments, and fossil content. However, many are fossiliferous. Many formations exposed within the park also crop out on private lands just outside of its boundaries and so some fossils from just outside the park are also included here as well. In general, older, Late Cretaceous open marine carbonate strata are separated by unconformities representing relatively brief geologic intervals. Younger, Late Cretaceous (marine shelf) strata generally grade conformably into, and sometimes inter-tongue with, broadly coeval terrestrial rocks. These deposits then grade conformably into overlying Paleocene strata. Some localized unconformities are present in some strata (e.g., the Aguja/Javelina formations contact) as the result of penecontemporaneous erosion (i.e., stream downcutting which occurred simultaneously with overbank deposition in some areas), but these minor depositional gaps generally do not represent geologically significant intervals. Significant erosion of Cretaceous and Paleocene strata did occur as the result of Laramide uplifting in Eocene time however these eroded deposits were then covered by even younger fluvial deposits, volcaniclastics, and extrusive rocks. Basin and range development along with continued erosion of the Chisos Mountains volcanic complex initiated yet another period of deposition which resulted in infilling of the surrounding bolson. Despite the presence of unconformities, many of the strata within the park and immediate surrounds preserve a relatively continuous, 135 million-year-long depositional sequence. The following formations are arranged in stratigraphic succession (low to high) (Fig.
Ordovician, marine, around 50 m thick. The Maravillas was deposited in a deep-water, basinal environment (
Silurian-Devonian, marine, only 20 m thick. The origin of both the novaculite and chert members leads to contrasting interpretations of water depth during deposition (e.g.,
Mississippian - Pennsylvanian, marine, variably thick from 15-200 m. Deep-water sediments, thin to thickly bedded sandstone and dark gray, brown, and black shale. Several small outcrops are situated in the northernmost part of BIBE. No fossils have been collected from the park however nearby areas have produced conodonts, foraminifera, and a few Pennsylvanian Period plant fossils (
Marine, massive, about 100-150 m thick. Primarily a massive limestone but contains clay, minor sandstone, marl, and conglomerate deposited in near-shore tidal and sub-tidal marine environs (
Marine, generally 20-45 m thick. Lagoonal sediments (
Marine, massive, from 100-150 m thick. Open lagoon, tidal flat, and rudistid biostromal facies (
Marine, around 25-30 m thick. Transgressional marine sediments containing shale, marl, thin nodular limestone ledges (
Marine, massive, up to 225 m thick. Open shelf carbonate environments (
Marine, fissile, around 1-35 m thick. A regressive marine environment facilitated development of this shaly, shallow-water facies (
Marine, 20-30 m thick. Shallow, inner-shelf environment. This formation primarily crops out in eastern, southern. and southwestern areas of the park such as Dog Canyon, Dagger Mountain, Mariscal Mountain, and Mesa de Anguila (
Marine, massive to shaley, from 220-245 m thick. Foraminiferal limestone and shale deposited in relatively shallow, open marine (platform) conditions (
Marine shelf, 70-200 m thick. Calcareous clay shale and chalky limestone with concretionary intervals. The Pen Formation was deposited upon a shallow marine shelf. This unit also includes a westerly-thinning wedge of dark gray marine shale within the overlying Aguja Formation (e.g.,
Originally named "Rattlesnake Beds" by
Some facies within these units are very fossiliferous while others are not. Plant fossils are locally abundant in the Aguja Formation. These usually include fossilized woods from conifers, palms (monocots), and flowering plants (dicots). Rarely, tree stumps are found upright, situated in their original growing positions. Fossil leaves have been found in a couple areas preserved as carbonate films within mudstone horizons or, in one area, as impressions within reworked volcanic ash. This ash bed and its fossils are currently under study by the author (S.W.). Aguja invertebrate fossils include bivalves, gastropods, cephalopods and rarely, crustaceans. Trace fossils from some of these taxa are also relatively common (e.g., Ophiomorpha burrows).
Occasionally, vertebrate fossils (and microfossils) are also found at various stratigraphic intervals in strata representing numerous environs. Taxa include sharks, fish, turtles, crocodilians, as well as dinosaurs among other reptiles. Very rarely, small fossil mammals are encountered (mostly teeth) as are dinosaur eggshell fragments. The vertebrate fossil assemblage of the Aguja Formation is the most inclusive of its kind reported from southernmost North America.
Continental, 100-190 m thick. The formation can be found along the flanks of the Chisos Mountains and is well exposed along the drainages of Tornillo, Terlingua, and Dawson creeks, as well as Rough Run (
Isolated, broken vertebrate fossils are somewhat common within scree along deflated surfaces atop fluvial sandstone hogbacks but are also found in-situ at local intervals within overbank mudstones. Vertebrate fossils include those from fish, turtles, pterosaurs, dinosaurs, and small mammals (represented mostly by teeth). Vertebrate fossils usually occur as isolated, fragmentary bones. However, a few dinosaur skeletons have been found partially articulated or with bones in close association. Although typically well-preserved, Javelina Formation fossils are seemingly not as numerous as those of the underlying Aguja Formation. As such, I surmise that the paralic Aguja environment favored a greater variety (and populations) of vertebrate species and/or the paralic environment was more conducive the burial and preservation of remains.
Continental, around 40 m thick (widely variable) (e.g.,
The Cretaceous-Paleogene (K-Pg) boundary is situated in the lower third of the Black Peaks Formation although its exact stratigraphic position remains obscure. It has been defined within a two-meter section near the Grapevine Hills (
Continental, up to 400 m thick (widely variable). The Black Peaks Formation straddles the K-Pg boundary. The Paleogene portion of the formation contains inland floodplain deposits with thick, fluvial sandstones. It is exposed near Dogie Mountain, Grapevine Hills, and Tornillo Flat (
Plant fossils (mostly conifers) are rarely found in the lower part of the formation but are more common higher in section. Two, closely-space stratigraphic intervals of very large fossil dicot logs (Paraphylanthoxylon) in the middle portion (Torrejonian-Tiffinian) of the Black Peaks section (informally called the "log jam sandstone") suggest the post K-Pg resurgence of trees during this time. This fossil log horizon is conspicuous in many areas of the park and is a useful stratigraphic marker (
Continental, varies from around 30 to 70 m in thickness (e.g.,
Continental (upland), up to 350 m thick. This formation is exposed in the north-central portion of BIBE especially on Tornillo Flat (
Vertebrate fossils are widespread within the formation in BIBE as well as areas northwest of the park in the Devil's Graveyard Formation which is temporally coeval with the Canoe Formation (e.g.,
Continental (upland), from 500-700 m thick. The Chisos Formation is exposed in many areas of BIBE along the flanks of the Chisos Mountains (
Please note that the Burro Mesa Formation is not considered valid by all researchers and so both are included together here. Continental (volcanic), from 300-500 m thick. These typically massive, volcanically-derived strata are exposed in the central and southwest portions of BIBE in the Chisos Mountains and near Burro Mesa. They contain lavas, flow breccias, conglomerates, tuff, and tuffaceous sediments from various localized eruptive events and are apparently non-fossiliferous.
Continental (bolson deposits), up to 300 m thick. The formation is exposed on the west side of BIBE near Castolon (
Continental (bolson deposits), up to 150 m thick. This formation is exposed in the east-central portion of BIBE near Banta Shut-In. These include pink fine-grained sandstone, siltstone and red mudstone which represent distal alluvial fan facies in the eastern half of BIBE (Estufa Bolson). Vertebrate fossils include amphibians, reptiles, and mammals (including those from canids, camels, and primitive horses). This formation is exposed in areas along Tornillo Creek that are not easy to reach and it is likely that its fossiliferous nature has yet to be fully realized.
Continental (bolson deposits), variably thick up to 300 m. These formations are exposed in the western portion of BIBE near Sotol Vista and along the flanks of Tornillo Creek east of Dugout Wells and consist mostly of bolson deposits. They were originally identified as 'older gravels' by
Thin alluvial gravels, sands, silts, caliche-cemented silts, small dune fields harboring a variety of localized cut-and-fill structures and small head-cutting drainages harboring a variety of finely to poorly sorted rock types. These thin deposits form desert pavement atop alluvial terrace remnants where aeolian erosion and sheet-wash have often removed finer sediments (
Numerous cliffside grottos can be also found throughout BIBE. Of interest is the discovery within one of these near Mule Ears Peaks of remains pertaining to California condors which no longer live in the Big Bend region. Whether these remains are truly fossils or not is debatable. However, they are estimated to be thousands of years old (
'Taxonomy' is the scientific study of naming, defining, and classifying groups of biological organisms based on shared or differing morphological characteristics. The following taxonomic lists were compiled from hundreds of reliable sources. These included peer-reviewed scientific reports, graduate-level academic studies (e.g., Ph.D. dissertations and Master's Theses), field trip guidebooks, scientific abstracts, as well as verifiable first-hand accounts (current research) reported to the author by qualified researchers. In the interest of compiling a comprehensive taxonomic catalog of fossils from Big Bend National Park (and immediate surrounds), all reported taxa are included. This distinction is important because, in some cases, a species reported decades ago may have more recently been taxonomically re-classified differently as something else. As a result, some older taxa may no longer be valid and/or a few may be recorded twice as the result of different taxonomic interpretations. In other cases, taxa may be listed multiple times with varying degrees of certainty (e.g., sometimes with a question mark or designated as a possible new species - see below). These are all included in the present report as they may represent more than one species. This circumstance serves to illustrate our constantly changing understanding of how species relate to one another.
The taxonomic lists presented here are organized alphabetically within classes of the Linnaean taxonomic classification system. Their common names are also provided as well. This serves to simplify the identification and listing of each species (from the perspective of interpretation) and allows for the convenient addition of future data within each table. This simplified method was chosen because taxonomic groupings at family-level (and below) often complicate matters to the point of utter confusion for non-scientists - especially as classification systems and taxonomic relationships are revised when new information comes to light.
Furthermore, additional taxa have been added to the original catalog produced by
These lists also embrace the 'morphotype concept' of taxonomy and is used so that scientists can communicate with each other more effectively. For example, different types of plant fossils from a single taxon are often named differently because that plant species may be expressed in the paleontological record by multiple fossil morphotypes (such as fossil wood, leaf impressions, and/or pollen). From this example, unless all three types of plant fossils are found in close association, each type of fossil cannot be conclusively determined to pertain to the same plant species. Hence, each form is given its own name until a direct association can be confirmed. As such, a single plant species may unknowingly be represented here by more than one morphotype (and scientific name). Also included in these lists are non-body fossils (such as crustacean burrows and dinosaur eggshell fragments) produced by a living organism. These are also classified and named using the morphotype concept since they do not represent the actual fossilized remains of a particular animal, but only the preserved evidence of its lifeway.
Also included are the formations in which the fossils occur as well as the original (or significant) publications in which they were reported. Because commonly encountered species (e.g., various sharks among others) are mentioned in numerous reports, it is simply impractical to include every reference for many of these commonly reported species. It is, therefore, up to the reader to use the listed sources as springboards for further research. Problematic taxa and /or references indicated by an asterisk are discussed at the bottom of each list.
Finally, the reader needs to be aware that the author of the present work did not make any of the taxonomic interpretations for a particular species listed herein unless (as in a few cases) he actually authored one of the referenced papers. Among the names of the species listed herein, the reader will sometimes see various abbreviations associated with them. The applications of abbreviations such as these are standard practice among taxonomists (e.g., see
Since 1907, when Johan Udden first reported the occurrence of fossil wood in what would become Big Bend National Park, over 300 fossil plant taxa have been described including flowering plants (dicots), palms (monocots), conifers, tree ferns, leaf impressions, algae, palynomorphs and tree resin (amber). Because of the changing environment over time, fossil plant remains range from marine, coastal, and inland varieties spanning a diverse range of paleohabitats. Numerous type specimens (nine) have been formally described with several others having been recognized but not yet reported. Two-thirds of the fossil plant species reported from BIBE pertain to palynomorphs (e.g., pollen, spores, fungi, etc.).
Although fragmentary fossil wood specimens are observed within many continental strata in BIBE, they are uncommon or absent in most locations. However, a few horizons produce spectacular fossil logs, sometimes by the dozens (
The degree of preservation involving fossil woods from BIBE ranges from those having experienced near-complete permineralization (i.e., exhibiting few visible diagnostic attributes) to those that preserve very detailed morphological features such as growth rings and cellular structure such as compression wood, parenchyma, and cross-field pitting (e.g.,
Fossil wood is widespread in BIBE and many specimens are situated near roads, trails or camping areas. From the public's perspective, they are also some of the most recognizable types of fossils in BIBE and as a result, are often reported to park management by visitors. Because they are somewhat obvious and popular, "petrified" woods also remain one of the most easily vandalized fossil types in many NPS fossil-parks which has led to the loss of valuable scientific information (
CLASSIFICATION | TAXON | FORMATIONS | REFERENCES | |||||||||||
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GR | TC | DC | SE | DR | BO | PN | AG | JV | BP | HH | CN | |||
CYANOPHYTA | ||||||||||||||
blue-green algae | Stromatolites | X | X | X |
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ULVOPHYCEAE | ||||||||||||||
calcareous algae | Cylindroporella sp. | X | X | X |
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Heteroporella sp. | X |
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Permocalculus irenae | X | X |
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Salpingoporella sp. | X |
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Terquemella sp. | X |
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BRYOPSIDOPHYCEAE | ||||||||||||||
calcareous algae | Boueina sp. | X |
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CHAROPHYTA | ||||||||||||||
algae (oogonia) | ?Charophytes Indet. | X | X |
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PHAEOPHYCEAE | ||||||||||||||
brown algae | Fucales indet. | X | X |
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Halymenites sp. | X |
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PTERIDOPSIDA | ||||||||||||||
tree ferns | Tempskya sp. | X |
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CONIFEROPHYTA | ||||||||||||||
GYMNOSPERM TREES | Abeitoxylon maxwellii Δ | X |
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Araucariaceae indet. | X | X | X |
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Araucarioxylon maxwellii Δ | X | * |
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Araucarioxylon sp. | X |
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Brachyphyllum sp. | X |
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Coniferophyta indet. | X |
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Glyptostrobus sp. | X |
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Podocarpaceae indet. | X | X |
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Sequoia sp. | X |
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?Sequoia reichenbachia | X |
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Thuyoxylon maxwellii Δ | X |
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Tornilloxylon maxwellii Δ | X |
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MONOCOTYLEDONEAE | ||||||||||||||
PALM TREES | Sabal bigbendense • | X |
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Sabal bracknellense | X |
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Sabalites ungeri (leaf impression) | X |
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DICOTYLEDONEAE | ||||||||||||||
ANGIOSPERM TREES | Acalyphoxylon maxwellii Δ | X |
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Agujoxylon olacaceoides • | X |
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Baasoxylon parenchymatosum • | X | X |
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Baileyan Big Bend wood type I (scrambling vine) | X |
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Bombacoxylon langstoni • | X |
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Canarioxylon maxwellii Δ | X |
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Chimarrhioxylon maxwellii Δ | X |
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Cissus sp. | X |
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Crataveoxylon maxwellii Δ | X |
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cf. Cunonioxylon sp. | X |
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Dialyantheroxylon maxwellii Δ | X |
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Dicotyledoneae indet. | X |
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Ericales indet. | X |
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Gassonoxylon araliosum • | X | X |
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Hasseltioxylon maxwellii Δ | X |
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Javelinoxylon multiporosum • | X |
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Metcalfeoxylon kirtlandense | X |
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Pachirioxylon maxwellii Δ | X |
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Pageoxylon cretaceum • | X |
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Paraphyllanthoxylon abbottii • | X |
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cf. Paraphyllanthoxylon anazasii | X | X |
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Platanoid wood type I (scrambling vine) | X |
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Platanoid wood type II (scrambling vine) | X |
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Platanoxylon sp. | X |
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cf. Platinus haydenii | X |
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Preplatanoxylon maxwellii Δ | X | * |
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Pycnanthoxylon maxwellii Δ | X |
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Sabinoxylon wicki • | X |
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Sloaneoxylon maxwellii Δ | X |
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Vitexoxylon maxwellii Δ | X |
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FOSSIL LEAVES | ||||||||||||||
VARIOUS TAXA | Ampelopis acerifolia | X | * |
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Carbonized leaf impressions | X | Montogomery and Clark 2016 | ||||||||||||
Cheirolepidaceae | X |
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Chloranthaceae | X |
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Ficus cf. F. tennesseensis | X | * |
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FOSSIL LEAVES (continued) | ||||||||||||||
VARIOUS TAXA | "Hamamelid-like" | X |
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Laurus socialis | X | * |
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Monocotyledonae indet. | X | Lehman and Langston unpublished | ||||||||||||
Paracredneria sp. | X |
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cf. Persea | X |
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Platanacea | X |
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Platanus raynoldsi | X | * |
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Podocarpaceae | X |
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Rhizophoracea | X |
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Taxodiaceae | X |
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Typha sp. | X | * |
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Dicot and fern leaf impressions in volcanic ash - multiple taxa under study | X | Wick in prep. | ||||||||||||
MISCELLANEOUS | ||||||||||||||
Fossilized tree resin (amber) | X |
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Indeterminate fossil wood | X |
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PALYNOMORPHS | ||||||||||||||
SPORES, POLLEN, FUNGI, ETC. | Alnipollenites trina | X | * |
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Alnipollenites verus | X | * |
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Alsophiliidites kerquelensis | X | * |
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Apiculatisporites sp. | X |
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Appendicisporites prolematicus | X |
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Appendicisporites sp. | X |
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Appendicisporites tricornitatus | X |
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Apteodinium sp. | X |
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Arecipites microreticulatus | X |
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Arecipites sp. | X |
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Baltisphaeridium sp. | X | X |
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Betulaceae indet. | X |
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Betulaceoipollenites infrequens | X | * |
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Bombacacopites nacimientoensis | X | * |
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Caligodinium sp. | X | X |
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Callialasporites sp. | X |
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Camarozonsporites rudis | X |
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Camarozonsporites sp. | X | X |
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Canningia sp. | X |
|
||||||||||||
Cannosphaeropsis sp. | X |
|
||||||||||||
Caryapollenites simplex | X |
|
||||||||||||
Caryapollenites sp. | X |
|
||||||||||||
Casaurinidites sp. | X |
|
||||||||||||
Ceratosporites sp. | X |
|
||||||||||||
Cerodinium diebelii | X | X |
|
|||||||||||
Chatangiella sp. | X |
|
||||||||||||
Cicatricosisporites sp. | X |
|
||||||||||||
Cicatricosporites dorogensis | X | * |
||||||||||||
Cingulatisporites sp. | X |
|
||||||||||||
Circulina parva | X | X |
|
|||||||||||
Classopollis classoides | X |
|
||||||||||||
Cleistosphaeridium polypes | X |
|
||||||||||||
Complexipollis abditus | X |
|
||||||||||||
Complexipollis sp. | X |
|
||||||||||||
Concavisporites cf. arugulatus | X |
|
||||||||||||
Cordosphaeridium sp. | X |
|
||||||||||||
Corsinipollenites sp. | X |
|
||||||||||||
Cupuliferoipollenites pusillus | X |
|
||||||||||||
Cupuliferoipollenites sp. | X |
|
||||||||||||
Cyathidites australis | X |
|
||||||||||||
Cyathidites foveolatus | X |
|
||||||||||||
Cyathidites minor | X |
|
||||||||||||
Cyathidites sp. | X |
|
||||||||||||
Cycadopites carpentieri | X |
|
||||||||||||
Cycadopites pollicularis | X | * |
||||||||||||
Cycadopites scabratus | X | X | X | * |
||||||||||
Cycadopites sp. | X | X |
|
|||||||||||
Cyclopsiella sp. | X | X |
|
|||||||||||
Cyrilla minima | X | * |
||||||||||||
Cyrillaceaepollenites exactus | X |
|
||||||||||||
Deflandrea cooksoniae | X |
|
||||||||||||
Deflandrea obscura | X | X |
|
|||||||||||
Deflandrea oebisfeldensis | X |
|
||||||||||||
Deflanrdea sp. | X | X |
|
|||||||||||
Deltoidospora diaphana | X |
|
||||||||||||
Deltoidospora mesozoica | X |
|
||||||||||||
PALYNOMORPHS (continued) | ||||||||||||||
SPORES, POLLEN, FUNGI, ETC. | Deltoidospora minor | X |
|
|||||||||||
Deltoidospora sp. | X |
|
||||||||||||
cf. Didymoporisporonites sp. | X |
|
||||||||||||
Dinogymnium sp. | X | X |
|
|||||||||||
Echinatisporites longechinus | X |
|
||||||||||||
Echinatisporites sp. | X | X |
|
|||||||||||
Engelhardtia microfoveolate | X | * |
||||||||||||
Equisetosporites multicostatus | X |
|
||||||||||||
Exesipollenites tumulus | X |
|
||||||||||||
Faguspollenites granulatus | X |
|
||||||||||||
cf. Foveodiporites sp. | X |
|
||||||||||||
Gleicheniidites senonicus | X | X | * |
|||||||||||
Gleicheniidites senonicus | X |
|
||||||||||||
Gleicheniidites sp. | X |
|
||||||||||||
Gnetaceapollenites eocenipites | X |
|
||||||||||||
Granulatisporites sp. | X |
|
||||||||||||
Gymnodinium sp. | X |
|
||||||||||||
Hymenozonotriletes sp. | X |
|
||||||||||||
Hyphites sp. | X |
|
||||||||||||
hypoxylonites sp. | X |
|
||||||||||||
Hystrichosphaera sp. | X |
|
||||||||||||
Hystrichosphaeridium sp. | X |
|
||||||||||||
Hystrichosphaeridium tubiferum | X |
|
||||||||||||
Inapertisporites sp. | X | X |
|
|||||||||||
Inaperturepollenites magnus | X |
|
||||||||||||
Inaperturepollenites sp. | X | X |
|
|||||||||||
Inaperturopollenites dubius | X | * |
||||||||||||
Interpollis supplingensis | X |
|
||||||||||||
Intertrilites scrobiculatus | X | * |
||||||||||||
Intratrioporopollenites sp. | X |
|
||||||||||||
Kuylisporites scutatus | X |
|
||||||||||||
Kuylisporites sp. | X |
|
||||||||||||
Lacrimasporites levis | X | * |
||||||||||||
Laevigatosporites haardti | X | * |
||||||||||||
Laevigatosporites ovatus | X | X | * |
|||||||||||
Laevigatosporites percrassus | X | * |
||||||||||||
Laevigatosporites sp. | X |
|
||||||||||||
Leiotriletes sp. | X |
|
||||||||||||
Leptodinium sp. | X |
|
||||||||||||
Lilacidites dividuus | X |
|
||||||||||||
Lilacidites leei | X |
|
||||||||||||
Lilacidites sp. | X | X |
|
|||||||||||
Lilacidites variegatus | X |
|
||||||||||||
Liliacidites cf. L. complexus | X | X |
|
|||||||||||
Liliacidites sp. | X | * |
||||||||||||
Lusatisporis indistincta | X |
|
||||||||||||
Lusatisporis sp. | X |
|
||||||||||||
Lycopodiumsporites sp. | X |
|
||||||||||||
Lygodiumsporites sp. | X |
|
||||||||||||
Margocolporites cribellatus | X |
|
||||||||||||
Margocolporites sp. | X |
|
||||||||||||
Matonisporites cf. M. phelbopteroides | X |
|
||||||||||||
Michrystridium sp. | X |
|
||||||||||||
Microretiulatasporites cf. M. uniformis | X |
|
||||||||||||
Microretiulatasporites sp. | X |
|
||||||||||||
Microthyrites sp. | X |
|
||||||||||||
Momipites cf. M. coryloides | X |
|
||||||||||||
Momipites cf. M. tenuipolis | X |
|
||||||||||||
Momipites cf. M. wyomingensis | X |
|
||||||||||||
Momipites coryloides | X | * |
||||||||||||
Momipites sp. | X | X |
|
|||||||||||
Monocolpopollenites cf. M. magnus | X |
|
||||||||||||
Monocolpopollenites sp. | X |
|
||||||||||||
Monoporisporites stoverii | X | * |
||||||||||||
Monosulcites cf. M. glottus | X |
|
||||||||||||
Monosulcites perispinosis | X |
|
||||||||||||
Monosulcites sp. | X | X | * |
|||||||||||
Multilinaenites sp. | X |
|
||||||||||||
Multiporopollenites sp. | X |
|
||||||||||||
Neoraistrickia sp. | X |
|
||||||||||||
Nyssapollenites analepticus | X |
|
||||||||||||
Nyssoidites larsoni | X | * |
||||||||||||
Osmundacidites cf. O. wellmanii | X |
|
||||||||||||
Osmundacidites sp. | X |
|
||||||||||||
PALYNOMORPHS (continued) | ||||||||||||||
SPORES, POLLEN, FUNGI, ETC. | Osmundacidites wellmanii | X |
|
|||||||||||
Ovoidites ligneolus | X |
|
||||||||||||
Palaeohystrichophora infusorioides | X |
|
||||||||||||
Palaeohystrichophora sp. | X |
|
||||||||||||
Paleostomocystis sp. | X |
|
||||||||||||
Palmaepollenites tranquilis | X |
|
||||||||||||
Palmaepollenites cf. P. tranquillis | X |
|
||||||||||||
Palmaepollenites sp. | X | X |
|
|||||||||||
Parvisacctes radiatus | X |
|
||||||||||||
Parvisacctes sp. | X |
|
||||||||||||
Pediastrum sp. | X |
|
||||||||||||
Peregrinipollis sp. | X |
|
||||||||||||
Phelodinium magnifica | X |
|
||||||||||||
Pinus haploxylon | X | * |
||||||||||||
Pinuspollenites sp. | X |
|
||||||||||||
Planctonites sp. | X |
|
||||||||||||
Plicapollis retusus | X |
|
||||||||||||
Plicapollis sp. | X |
|
||||||||||||
Plicatopollis cf. C. plicata | X |
|
||||||||||||
Pluricellaesporites sp. | X |
|
||||||||||||
Podocarpidites sp. | X |
|
||||||||||||
Polyadosporites sp. | X |
|
||||||||||||
Polycingulatisporites reduncus | X |
|
||||||||||||
Portalites sp. | X |
|
||||||||||||
Proteacidites marginus | X |
|
||||||||||||
Proteacidites molis | X |
|
||||||||||||
Proteacidites retusus | X |
|
||||||||||||
Proteacidites sp. | X |
|
||||||||||||
Proteacidites thalmanni | X |
|
||||||||||||
Pseudolasopollis sp. | X |
|
||||||||||||
Pseudolasopollis ventosa | X |
|
||||||||||||
Psilatricolporites sp. | X |
|
||||||||||||
Psilatriletes sp. | X |
|
||||||||||||
Punctatosporites major | X |
|
||||||||||||
Punctatosporites sp. | X | * |
||||||||||||
Rectosulcites latus | X | X |
|
|||||||||||
Reticulatosporites sp. | X | X | * |
|||||||||||
Retipollenites cf. R. confusus | X |
|
||||||||||||
Retipollenites sp. | X | X |
|
|||||||||||
Retitricolpites florentinus | X |
|
||||||||||||
Retitriletes muricatus | X |
|
||||||||||||
Retitriletes sp. | X |
|
||||||||||||
Retricolpites sp. | X |
|
||||||||||||
Rhiopites globosus | X | * |
||||||||||||
Rhoipites cf. R. cryptoporus | X |
|
||||||||||||
Rhoipites sp. | X |
|
||||||||||||
Rugulatisporites quintus | X | * |
||||||||||||
Sabalpollenites cf. convexus | X |
|
||||||||||||
Scabritricolpites sp. | X |
|
||||||||||||
Schizaeoisporites eocaenicus | X |
|
||||||||||||
Schizoporis sp. | X | * |
||||||||||||
Schizosporis parvus | X |
|
||||||||||||
Schizosporis sp. | X |
|
||||||||||||
Seductisporites sp. | X |
|
||||||||||||
Septohyphites sp. | X |
|
||||||||||||
Sphagnumsporites antiquasporites | X |
|
||||||||||||
Sphagnumsporites sp. | X |
|
||||||||||||
Spinidinium densispenatum | X | X |
|
|||||||||||
Spinidinium microceratum | X |
|
||||||||||||
Spinidinium sp. | X | X |
|
|||||||||||
Staphlosporonites sp. | X |
|
||||||||||||
Stereisporites cf. S. crassus | X |
|
||||||||||||
Stereisporites psilatus | X | * |
||||||||||||
Stereisporites sp. | X | X |
|
|||||||||||
Striadiporites sp. | X |
|
||||||||||||
Subtilisphaera sp. | X | X |
|
|||||||||||
Subtrudopollis sp. | X |
|
||||||||||||
Syncolporopollenites sp. | X |
|
||||||||||||
Taurocusporites cf. T. segmentatus | X |
|
||||||||||||
Taurocusporites sp. | X |
|
||||||||||||
Taxodiaceaepollenites hiatus | X | X | X | * |
||||||||||
Tetracellites sp. | X |
|
||||||||||||
Tetracolporopollenites manifestus | X | * |
||||||||||||
?Tetradites sp. | X |
|
||||||||||||
PALYNOMORPHS (continued) | ||||||||||||||
SPORES, POLLEN, FUNGI, ETC. | ?Tilia sp. | X |
|
|||||||||||
Todisporites minor | X |
|
||||||||||||
Triatriopollenites cf. T. pseudogranulatus | X |
|
||||||||||||
Triatriopollenites cf. T. pseudovestibulum | X |
|
||||||||||||
Triatripollenites rurensis | X |
|
||||||||||||
Triatripollenites sp. | X |
|
||||||||||||
Tricolpites cf. T. reticulatus | X |
|
||||||||||||
Tricolpites parvistriatus | X | * |
||||||||||||
Tricolpites sp. | X | X | * |
|||||||||||
Tricolpopollenites levitas | X |
|
||||||||||||
Tricolpopollenites microhenrici | X |
|
||||||||||||
Tricolpopollenites micropunctatus | X |
|
||||||||||||
Tricolpopollenites sp. | X |
|
||||||||||||
Tricolporites rhomboides | X |
|
||||||||||||
Tricolporites sp. | X |
|
||||||||||||
Tricolporopollenites kruschii | X |
|
||||||||||||
Tricolporopollenites cf. T. desultorius | X |
|
||||||||||||
Tricolporopollenites sp. | X |
|
||||||||||||
Tricolporopollenites triangulus | X |
|
||||||||||||
Triplanosporites pseudosinosus | X | * |
||||||||||||
Triplanosporites sinosus | X | * |
||||||||||||
Triporoletes novmexicanus | X |
|
||||||||||||
Triporopollenites bituitus | X |
|
||||||||||||
Triporopollenites coryloides | X | X |
|
|||||||||||
Triporopollenites robustus | X |
|
||||||||||||
Triporopollenites rugatus | X | * |
||||||||||||
Triporopollenites sp. | X | X |
|
|||||||||||
Trithyrodinium sp. | X | X |
|
|||||||||||
Trivestibulopollenites sp. | X |
|
||||||||||||
Trudopollis sp. | X |
|
||||||||||||
Ulmoideipites krempii | X |
|
||||||||||||
Undulatisporites sp. | X |
|
||||||||||||
Verrucatosporites sp. | X |
|
||||||||||||
Verrucingulatisporites sp. | X |
|
||||||||||||
Verrutriporites sp. | X |
|
||||||||||||
Vitipites affluens | X |
|
||||||||||||
Vitipites sp. | X |
|
||||||||||||
Wilsonipites sp. | X | X |
|
|||||||||||
Zlivisporis novamexicanum | X |
|
||||||||||||
Zlivisporis sp. | X |
|
||||||||||||
Zygnema sp. | X |
|
||||||||||||
HOLOTYPE = • | Glen Rose Limestone | Telephone Canyon Formation | Del Carmine Formation | Santa Elena Limestone | Del Rio Clay | Boquillas Formation | Pen Formation | Aguja Formation | Javelina Formation | Black Peaks Formation (K/Pg undifferentiated) | Hannold Hill Formation | Canoe Formation | ||
UNPUBLISHED NEW TAXON = Δ | ||||||||||||||
UNNAMED NEW TAXON = ο | ||||||||||||||
Lower Cretaceous | Upper Cretaceous | Cenozoic | ||||||||||||
FORMATIONS | ||||||||||||||
MISCELLANEOUS | Unidentified log fragments | Chisos Formation (Eocene) |
|
Over 500 fossil invertebrate taxa have been reported from BIBE including sponges, corals, bivalves, gastropods, ammonites, nautiloids and crustaceans, as well as a host of foraminifera. Invertebrates have been observed in many formations within BIBE from marine, brackish and freshwater facies. Five, scientifically valid type specimens have been described from BIBE and several 'new' taxa have yet to be formally reported.
Invertebrate fossils are regularly discovered in marine and brackish water facies within BIBE. Because of their abundance, form, and common occurrence along the modern shores of North America, invertebrate fossils are very popular as they are easily recognizable to park visitors of all ages. Fresh water taxa are much less common than their saltwater counterparts however they are occasionally discovered in lacustrine and fluvial deposits within some continental strata in the park. Many invertebrate fossils are preserved as steinkerns which represent the fossilized fill of a hollow organic structure (such as a mollusk shell) that formed when mud or sediment consolidated within the structure and the structure itself disintegrated or dissolved. Many invertebrate fossils are found in Lower Cretaceous, marine carbonate rocks along the fault-scarps which flank the northeastern and southwestern margins of the park. Upper Cretaceous forms are commonly preserved along with the remains of vertebrate taxa (such as sharks and mosasaurs) in near-shore marine mudstones and marls of the Boquillas and Pen formations surrounding the Chisos Mountains and exposed just west of the park. Some invertebrates from BIBE are particularly useful as stratigraphic index fossils. These include the ammonite Allocrioceras hazzardi (Young) and the bivalve Inoceramus undulatoplicatus (Roemer) both from the shallow marine, Boquillas Formation (e.g.,
The preservation of invertebrate fossils varies by formation and facies. Lower Cretaceous, marine carbonate rocks often preserve invertebrate fossils such as bivalves and gastropods, but these are often entombed in dense carbonate matrix and are very difficult to extract without damage. Microinvertebrates are also difficult to separate from these rocks and require laboratory preparation (thin section samples) to study the fossils within. Upper Cretaceous strata have produced well preserved, intact invertebrate specimens (e.g., ammonites and bivalves) which occur as steinkerns in carbonate facies or within concretionary horizons. Some are difficult to remove from bedrock while others can be quarried easily. Other bottom-dwelling invertebrates often occur in marine mudstones and shales as isolated individuals or in loose, congregated groups such as the oyster Flemingostrea pratti (Stephenson) and sea urchin Hemisaster (Desor). Occurrences of this type are often observed in horizons within marine or brackish water facies which may contain dozens of individuals which inhabited muddy estuarial bottoms. Some fossils, such as the oyster Crassostrea cusetta (Sohl and Kauffman) are often found in dense groups (formerly bioherms). Many individuals exhibit obvious warping of their shells as the result of a congested colonial lifeway. Trace fossils (e.g., burrow structures) are also routinely observed in various marine strata.
CLASS | TAXON | FORMATIONS | REFERENCES | |||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
GR | TC | DC | SP | SE | DR | BU | BO | PN | AG | JV | BP | |||
PROTOZOA | ||||||||||||||
single-celled org. | Spironema sp. | X |
|
|||||||||||
FORAMINIFERA | ||||||||||||||
PLANKTONIC CREATURES | Ammobaculites cuyleri | X |
|
|||||||||||
Ammobaculites dentonensis | X |
|
||||||||||||
Ammobaculites fragmentarium | X |
|
||||||||||||
Ammobaculites subcretacea | X | X |
|
|||||||||||
Anomalina plummerae | X |
|
||||||||||||
Archaeoglobigerina blowi | X | X | Graham 1995; |
|||||||||||
Archaeoglobigerina bosquensis | X | X | Graham 1995; |
|||||||||||
Archaeoglobigerina cretacea | X | X | Graham 1995 | |||||||||||
Bolivina textularoides | X |
|
||||||||||||
Bolivinita planata | X |
|
||||||||||||
Bolivinitella eleyi | X |
|
||||||||||||
Brachycythere sphenoides | X |
|
||||||||||||
Bulimina nannina | X |
|
||||||||||||
Buliminella carseyae | X |
|
||||||||||||
Buliminella cushman | X |
|
||||||||||||
Charentia sp. | X |
|
||||||||||||
Chrysalogonium cf. C. texanum | X |
|
||||||||||||
Citharina complanata | X |
|
||||||||||||
Coskinolinoides texanus | X | X | X |
|
||||||||||
Costallagerina thompsoni | X | X | Graham 1995 | |||||||||||
Costellagerina bulbosa | X | X | Graham 1995 | |||||||||||
Costellagerina phlegeri | X | X | Graham 1995 | |||||||||||
Costellagerina smithi • | X | Graham 1995 | ||||||||||||
Cribratina texana | X |
|
||||||||||||
Cuneolina cf. pavonia | X | X | X | X | X | X |
|
|||||||
Cythereis bicornis | X |
|
||||||||||||
Cythereis cf. C. austinensis | X |
|
||||||||||||
Cythereis dallasensis | X |
|
||||||||||||
Cytherella austinensis | X |
|
||||||||||||
Dentalina communis | X |
|
||||||||||||
Dentalina crypta | X |
|
||||||||||||
Dentalina debilis | X |
|
||||||||||||
Dentalina gracilis | X |
|
||||||||||||
Dentalina intrasegma | X |
|
||||||||||||
Dentalinopsis excavata | X |
|
||||||||||||
Dentalinopsis tricarinatum | X |
|
||||||||||||
Dentelina soluta | X |
|
||||||||||||
Dentilina hammensis | X |
|
||||||||||||
Dicarinella algeriana | X |
|
||||||||||||
Dicarinella asymetrica | X | X | Graham 1995 | |||||||||||
Dicarinella concavata | X | X | Graham 1995; |
|||||||||||
Dicarinella daileyi • | X | Graham 1995 | ||||||||||||
Dicarinella difformis | X | X | Graham 1995 | |||||||||||
Dicarinella indica | X | X | Graham 1995 | |||||||||||
Dictyoconus walnutensis | X | X | X |
|
||||||||||
Discorbis minima | X |
|
||||||||||||
Discorbis minutissima | X |
|
||||||||||||
Dorothia cf. D. alexanderi | X |
|
||||||||||||
Dorothia cf. D. bulletta | X |
|
||||||||||||
Dorothia stephensoni | X | X |
|
|||||||||||
Ellipsoidella gracillima | X |
|
||||||||||||
Eouvigerina plummerae | X |
|
||||||||||||
Favusella washitensis | X | X | X | X | X |
|
||||||||
Flabellammina clava | X |
|
||||||||||||
Frondicularia cordata | X |
|
||||||||||||
Gaudryina austinana | X | X |
|
|||||||||||
Gaudryina rudita | X | X |
|
|||||||||||
Globigerina cretacea | X |
|
||||||||||||
Globigerina rugosa | X |
|
||||||||||||
Globigerina saratogaensis | X |
|
||||||||||||
Globigerina sp. | X |
|
||||||||||||
Globigerina voluta | X |
|
||||||||||||
Globigerinella aissan a | X | X |
|
|||||||||||
Globigerinelloides asperus | X | X | Graham 1995 | |||||||||||
Globigerinelloides bentonensis | X |
|
||||||||||||
Globigerellinoides caseyi | X |
|
||||||||||||
FORAMINIFERA (continued) | ||||||||||||||
PLANKTONIC CREATURES | Globigerinelloides multispina | X | Graham 1995 | |||||||||||
Globiginerelloides prairiehillensis | X | Graham 1995 | ||||||||||||
Globoratalites umbilicatus | X |
|
||||||||||||
Globorotalia arca | X |
|
||||||||||||
Globorotalia cushmani | X |
|
||||||||||||
Globorotalia membranacea | X |
|
||||||||||||
Globorotalia sp. | X |
|
||||||||||||
Globotruncana arca | X |
|
||||||||||||
Globotruncana bulloides | X | Graham 1995 | ||||||||||||
Globotruncana canaliculata | X |
|
||||||||||||
Globotruncana contusa | X |
|
||||||||||||
Globotruncana cretacea | X |
|
||||||||||||
Globotruncana fornicata | X | X |
|
|||||||||||
Globotruncana lapparenti | X | Graham 1995 | ||||||||||||
Globotruncana marginata | X |
|
||||||||||||
Globotruncana membanacea | X |
|
||||||||||||
Globulina exerta | X |
|
||||||||||||
Globulina lacrima | X |
|
||||||||||||
Guembelitria graysonensis | X |
|
||||||||||||
Guembelitria harrisi | X |
|
||||||||||||
Gumbelina globocarinata | X |
|
||||||||||||
Gumbelina moremani | X |
|
||||||||||||
Gumbelina nuttalli | X |
|
||||||||||||
Gumbelina plummerae | X |
|
||||||||||||
Gumbelina pseudotessera | X |
|
||||||||||||
Gumbelina reussi | X |
|
||||||||||||
Gumbelina striata | X |
|
||||||||||||
Gyroidina depressa | X | X |
|
|||||||||||
Gyroidina girardana | X |
|
||||||||||||
Gyroidina globosa | X | X |
|
|||||||||||
Haplofragmoides concava | X |
|
||||||||||||
Haplostiche texana | X |
|
||||||||||||
Hastigerinella alexanderi | X |
|
||||||||||||
Hastigerinella moremani | X |
|
||||||||||||
Hastigerinella simplex | X |
|
||||||||||||
Hastigerinella watersi | X |
|
||||||||||||
Hastingerinoides alexanderi | X | X | Graham 1995 | |||||||||||
Hastingerinoides watersi | X | X | Graham 1995 | |||||||||||
Hedbergella delrioensis | X |
|
||||||||||||
Hedbergella planispira | X | X |
|
|||||||||||
Hedbergella simplex | X |
|
||||||||||||
Hedbergella sp. | X | X | X | X | X |
|
||||||||
Heterohelix cf. moremani | X | X | X | X |
|
|||||||||
Heterohelix globulosa | X | X | Graham 1995; |
|||||||||||
Heterohelix moremani | X | X |
|
|||||||||||
Heterohelix pulchra | X | Graham 1995 | ||||||||||||
Heterohelix reussi | X | X |
|
|||||||||||
Heterohelix semicostata | X | Graham 1995 | ||||||||||||
Heterohelix striata | X | Graham 1995 | ||||||||||||
Kyphopyxa christneri | X | X |
|
|||||||||||
Lagena apiculata | X |
|
||||||||||||
Lagena hispida | X |
|
||||||||||||
Lagena striatifera | X |
|
||||||||||||
Lagena sulcata | X |
|
||||||||||||
Lenticulina gaultina | X |
|
||||||||||||
Lenticulina rotulata | X | X |
|
|||||||||||
Lenticulina sp. | X | X | X | X |
|
|||||||||
Lingulina lamellata | X |
|
||||||||||||
Lingulina nodosaria | X |
|
||||||||||||
Lingulina sp. | X |
|
||||||||||||
Loxostumum cushmani | X |
|
||||||||||||
Marginotruncana angusticarenata | X | X | Graham 1995 | |||||||||||
Marginotruncana bigbendensi s | X | Graham 1995 | ||||||||||||
Marginotruncana coronat a | X | X | Graham 1995; |
|||||||||||
Marginotruncana marginata | X | X | Graham 1995; |
|||||||||||
Marginotruncana pseudolinneian a | X | X | Graham 1995 | |||||||||||
Marginotruncana renzi | X | X | Graham 1995 | |||||||||||
Marginotruncana undulata | X | X | Graham 1995 | |||||||||||
Marginulina austinana | X | X |
|
|||||||||||
Marginulina directa | X |
|
||||||||||||
Marssonella oxycona | X | X |
|
|||||||||||
FORAMINIFERA (continued) | ||||||||||||||
PLANKTONIC CREATURES | Neobulimina canadensis | X |
|
|||||||||||
Neobulimina irregularis | X |
|
||||||||||||
Neobulimina minima | X |
|
||||||||||||
Neoflabellina cushmani | X | X |
|
|||||||||||
Neoflabellina hebronensis | X |
|
||||||||||||
Neoflabellina suturalis | X |
|
||||||||||||
Nezzezata conica | X |
|
||||||||||||
Nezzezata simplex | X |
|
||||||||||||
Nodosaria affinis | X | X |
|
|||||||||||
Nodosaria barkeri | X |
|
||||||||||||
Nodosaria brandi | X |
|
||||||||||||
Nodosaria distans | X |
|
||||||||||||
Nodosaria obscura | X |
|
||||||||||||
Nodosaria scotti | X |
|
||||||||||||
Nodosaria sp. | X |
|
||||||||||||
Nodosaria tappanae | X |
|
||||||||||||
Opthalmidium sp. | X | X | X |
|
||||||||||
Orbitolina texana | X |
|
||||||||||||
Ovalviolina cf. maccognoae | X |
|
||||||||||||
Palmula pilulata | X | X |
|
|||||||||||
Paracypris angusta | X |
|
||||||||||||
Parathalmanniella appeninnica | X |
|
||||||||||||
Patellina subcretacea | X |
|
||||||||||||
Peneroplis sp. | X |
|
||||||||||||
Planomalina sp. | X |
|
||||||||||||
Planularia cf. P. dissona | X |
|
||||||||||||
Planulina arimensis | X |
|
||||||||||||
Planulina austinana | X | X |
|
|||||||||||
Planulina eaglefordensis | X |
|
||||||||||||
Planulina kansasensis | X |
|
||||||||||||
Pleurostomella austiniana | X |
|
||||||||||||
Pleurostomella watersi | X |
|
||||||||||||
Praeglobotruncana delrioensis | X |
|
||||||||||||
Praeglobotruncana stephani | X |
|
||||||||||||
Praeglobotruncana sp. | X |
|
||||||||||||
Pseudofrondicularia undulosa | X | X |
|
|||||||||||
Pseudoguembelina pessagnoi • | X | Graham 1995 | ||||||||||||
Pseudoguembelina halesi • | X | X | Graham 1995 | |||||||||||
Pseudotextularia elongata | X | X | Graham 1995 | |||||||||||
Pterygocythere cf. P. saratogana | X |
|
||||||||||||
Pyrulina cylindroides | X |
|
||||||||||||
Quinqueloculina aeschria | X |
|
||||||||||||
Rectogumbelina texana | X |
|
||||||||||||
Reophax difflugiformis | X |
|
||||||||||||
Reophax sp. | X |
|
||||||||||||
Robulus munsteri | X |
|
||||||||||||
Robulus taylorensis | X |
|
||||||||||||
Rosita fornicata | X |
|
||||||||||||
Rotilapora montsavensis | X |
|
||||||||||||
Rotalipora sp. | X | X |
|
|||||||||||
Saracenaria triangularis | X |
|
||||||||||||
Schackoina multispinata | X | X | Graham 1995 | |||||||||||
Scherochorella sp. | X |
|
||||||||||||
Sculptobaculites goodlandensis | X | X | X | X |
|
|||||||||
Shakoina sp. | X |
|
||||||||||||
Siderolites sp. | X |
|
||||||||||||
Sigalia alpina | X | Graham 1995 | ||||||||||||
Sigalia deflaensis | X | Graham 1995 | ||||||||||||
Spirillina minima | X |
|
||||||||||||
Spiroplectammina laevis | X |
|
||||||||||||
Spiroplectammina lalickeri | X |
|
||||||||||||
Spiroplectammina longa | X |
|
||||||||||||
Spiroplectammina nuda | X |
|
||||||||||||
Textularia rioensis | X |
|
||||||||||||
Textularia sp. | X |
|
||||||||||||
Thalmaninnella brotzeni | X |
|
||||||||||||
Thomasinella sp. | X |
|
||||||||||||
Ticinella sp. | X |
|
||||||||||||
Valvulinaria loetterei | X |
|
||||||||||||
Ventilabrella austiniana | X | X |
|
|||||||||||
Ventilabrella cf. V. browni | X | Graham 1995 | ||||||||||||
FORAMINIFERA (continued) | ||||||||||||||
PLANKTONIC CREATURES | Ventilabrella glabrata | X | Graham 1995 | |||||||||||
Virgulina tegulata | X |
|
||||||||||||
Vitriwebbina biosculata | X |
|
||||||||||||
Washitella typica | X |
|
||||||||||||
Whiteinella aprica | X | X | Graham 1995 | |||||||||||
Whiteinella archaeocretacea | X | X | Graham 1995 | |||||||||||
SPONGIAE | ||||||||||||||
sponges | Cliona sp. | X | X | X |
|
|||||||||
Myliusia sp. | X |
|
||||||||||||
ARTICULATA | ||||||||||||||
brachiopods | Kingena wacoensis | X |
|
|||||||||||
Terabratulina brewsterensi s | X | X |
|
|||||||||||
Terabratulina sp. | X |
|
||||||||||||
STENOLAEMATA | ||||||||||||||
bryozoans | Cyclostomata indet. | X |
|
|||||||||||
POLYCHAETA | ||||||||||||||
annelid worms | Hamulus onyx | X |
|
|||||||||||
Serpula cf. S. adnata | X |
|
||||||||||||
Serpula cretacea | X | X |
|
|||||||||||
ANTHOZOA | ||||||||||||||
stoney corals | Faviidae indet. | X |
|
|||||||||||
ECHINOIDEA | ||||||||||||||
SEA URCHINS | Echinoidia indet. | X |
|
|||||||||||
Enallaster calvini | X | X |
|
|||||||||||
Enallaster inflatus | X |
|
||||||||||||
Enallaster mexicanus | X |
|
||||||||||||
Enallaster sp. | X |
|
||||||||||||
Enallaster texana | X |
|
||||||||||||
Hemiaster calvini | X |
|
||||||||||||
Hemiaster sp. | X |
|
||||||||||||
Hemiaster texanus | X |
|
||||||||||||
Hemiasteridae indet. | X |
|
||||||||||||
Heteraster sp. | X | X |
|
|||||||||||
Heteraster texana | X |
|
||||||||||||
Heterohelix globulosa | X |
|
||||||||||||
Heterohelix moremani |
|
|||||||||||||
Heterohelyx sp. | X |
|
||||||||||||
Holectypus limitus | X |
|
||||||||||||
Holectypus sp. | X |
|
||||||||||||
Leiotomaster bosei | X |
|
||||||||||||
Macraster sp. | X |
|
||||||||||||
Phymosoma sp. | X |
|
||||||||||||
Pseudodiadema sp. | X |
|
||||||||||||
CRINOIDA | Saccocoma sp. | X |
|
|||||||||||
BIVALVIA | ||||||||||||||
CLAMS, OYSTERS, MUSSELS | Amphidonte sp. | X | X |
|
||||||||||
Anomia cf. A. mexicana | X |
|
||||||||||||
Anomia cf. A. argentaria | X |
|
||||||||||||
Anomia cf. A. tellinoides | X |
|
||||||||||||
Aphrodina sp. | X |
|
||||||||||||
Aphrodina tippana | X | X |
|
|||||||||||
Arcidae indet. | X |
|
||||||||||||
Arcopagella sp. | X |
|
||||||||||||
Astartidae indet. | X |
|
||||||||||||
Bivalvia indet. | X |
|
||||||||||||
Brachiodontes sp. | X |
|
||||||||||||
Brachymeris alta | X |
|
||||||||||||
Callista sp. | X |
|
||||||||||||
Camptonectes burlingtonensis | X |
|
||||||||||||
Camptonectes sp. | X |
|
||||||||||||
Caprinidae indet. | X |
|
||||||||||||
Cardium carolinensis | X | ? |
||||||||||||
Cardium congestum | X |
|
||||||||||||
Cardium longstreeti | X | X |
|
|||||||||||
Cardium sp. | X | X | X | X |
|
|||||||||
Cardium subcongesta | X |
|
||||||||||||
Cardium vaughni | X |
|
||||||||||||
Chondrodontidae indet. | X |
|
||||||||||||
Cladoceramus undulotplicatus | X |
|
||||||||||||
Cladoceramus undulatoplicatus michaeli | X |
|
||||||||||||
BIVALVIA (Continued) | ||||||||||||||
CLAMS, OYSTERS, MUSSELS | Corbicula cytheriformis | X |
|
|||||||||||
Corbicula sp. | X |
|
||||||||||||
Corbula sp. | X |
|
||||||||||||
Cordiceramus sp. | X |
|
||||||||||||
Crassatella cf. C. vadosa | X |
|
||||||||||||
Crassatella cf. C. obliquata | X |
|
||||||||||||
Crassostrea cusseta | X |
|
||||||||||||
Crassostrea trigonalis | X |
|
||||||||||||
Cremnoceramus crassus crassus | X |
|
||||||||||||
Cremnoceramus deformis erectus | X |
|
||||||||||||
Cremnoceramus walterdorfensis | X |
|
||||||||||||
Cyclorisma carolinensis | X |
|
||||||||||||
Cyclorisma sp. | X |
|
||||||||||||
Cymbophora berryi | X | X |
|
|||||||||||
Cymbophora scabellum | X |
|
||||||||||||
Cymbophora sp. | X |
|
||||||||||||
Cymbophora trigonalis | X |
|
||||||||||||
Cymella bella | X |
|
||||||||||||
Cyprimera depressa | X |
|
||||||||||||
Cyprimeria gabbi | X | X |
|
|||||||||||
Cyprimeria roddai | X |
|
||||||||||||
Cyprimeria sp. | X | X |
|
|||||||||||
Cyprimeria texana | X |
|
||||||||||||
Cyprina sp. | X |
|
||||||||||||
Dianchora cf. austinensis | X |
|
||||||||||||
Didymotis costatus | X |
|
||||||||||||
Dreissena tippana | X |
|
||||||||||||
Durania austinensis | X |
|
||||||||||||
Durania sp. | X |
|
||||||||||||
Durania terlinguae | X | X | X |
|
||||||||||
Eoradiolites cf. E. davidsoni | X |
|
||||||||||||
Eoradiolites cf. E. quadratus | X |
|
||||||||||||
Etea sp. | X |
|
||||||||||||
Ethmocardium cf. E. welleri | X |
|
||||||||||||
Exogyra arietina | X |
|
||||||||||||
Exogyra cancellata | X |
|
||||||||||||
Exogyra cartledgei | X |
|
||||||||||||
Exogyra clarki | X |
|
||||||||||||
Exogyra costata | X |
|
||||||||||||
Exogyra costata spinosa | X |
|
||||||||||||
Exogyra laeviuscula | X |
|
||||||||||||
Exogyra ponderosa ponderosa | X | X |
|
|||||||||||
Exogyra ponderosa whitneyi | X |
|
||||||||||||
Exogyra ponderosa erraticostata | X |
|
||||||||||||
Exogyra ponderosa upatoiensis | X |
|
||||||||||||
Exogyra quitmanensis | X |
|
||||||||||||
Exogyra sp. | X |
|
||||||||||||
Exogyra texana | X | X | X | X |
|
|||||||||
Exogyra whitneyi | X |
|
||||||||||||
Flemingostrea pratti | X |
|
||||||||||||
Flemingostrea subspatulata | X |
|
||||||||||||
Flemingostrea subspatulata n. subsp. + | X |
|
||||||||||||
Gastrochaena sp. | X |
|
||||||||||||
Granocardium sp. | X |
|
||||||||||||
Gryphaea cf. G. navia | X |
|
||||||||||||
Gryphaea corrugata | X |
|
||||||||||||
Gryphaea graysonana | X | X |
|
|||||||||||
Gryphaea mucronata | X | X | X |
|
||||||||||
Gryphaea sp. | X | X | X | X | X |
|
||||||||
Haploscapha grandis | X |
|
||||||||||||
Homomya sp. | X |
|
||||||||||||
Homomya washita | X |
|
||||||||||||
Ilmatogyra africana | X |
|
||||||||||||
Inoceramus anomalus | X |
|
||||||||||||
Inoceramus annulatus | X |
|
||||||||||||
Inoceramus arvanus | X |
|
||||||||||||
Inoceramus barabini | X |
|
||||||||||||
Inoceramus biconstrictus | X |
|
||||||||||||
Inoceramus cf. I. concentricus | X |
|
||||||||||||
Inoceramus cf. I. subquadratus | X |
|
||||||||||||
Inoceramus cumminsi | X |
|
||||||||||||
BIVALVIA (Continued) | ||||||||||||||
CLAMS, OYSTERS, MUSSELS | Inoceramus dakotensis | X |
|
|||||||||||
Inoceramus digitatus | X | Eley 1939 | ||||||||||||
Inoceramus exogyroides | X |
|
||||||||||||
Inoceramus ginterensis | X |
|
||||||||||||
Inoceramus (Haploscapha) grandis | X |
|
||||||||||||
Inoceramus howelli | X |
|
||||||||||||
Inoceramus labiatus | X | X |
|
|||||||||||
Inoceramus oblongus | X |
|
||||||||||||
Inoceramus perplexus | X |
|
||||||||||||
Inoceramus pertenuis | X |
|
||||||||||||
Inoceramus pictus | X |
|
||||||||||||
Inoceramus platinus | X |
|
||||||||||||
Inoceramus prefragilis | X |
|
||||||||||||
Inoceramus problematicus | X | X |
|
|||||||||||
Inoceramus rutherfordi | X |
|
||||||||||||
Inoceramus sp. | X | X | X |
|
||||||||||
Inoceramus umbonatus | X |
|
||||||||||||
Inoceramus undabundus | X |
|
||||||||||||
Inoceramus undulatoplicatus | X |
|
||||||||||||
Inoperna sp. | X |
|
||||||||||||
Isocardia medialis | X |
|
||||||||||||
Leda sp. | X |
|
||||||||||||
Leptosolen cf. L. quadrilaterus | X |
|
||||||||||||
Lima reticulata | X |
|
||||||||||||
Lima coahuilensis | X |
|
||||||||||||
Lima shumardi | X |
|
||||||||||||
Lima sp. | X |
|
||||||||||||
Lima wacoensis | X | X |
|
|||||||||||
Linearea sp. | X | X |
|
|||||||||||
Lingula cf. L. rauliniana | X |
|
||||||||||||
Lingula subspatulata | X |
|
||||||||||||
Liopistha bella | X |
|
||||||||||||
Lopha sp. | X |
|
||||||||||||
Lophia subovata | X |
|
||||||||||||
Lucina linearia | X |
|
||||||||||||
Lucina sp. | X |
|
||||||||||||
Lucinidae indet. | X |
|
||||||||||||
Lucinoma sp. | X |
|
||||||||||||
Lycettia tippanus | X |
|
||||||||||||
Mactra texana | X |
|
||||||||||||
Magadiceramus complicatus | X |
|
||||||||||||
Magadiceramus crenelatus | X |
|
||||||||||||
Magadiceramus subquadratus | X |
|
||||||||||||
Monopleuridae indet. | X |
|
||||||||||||
Mytiloides goppelnensis | X |
|
||||||||||||
Mytiloides herbichi | X |
|
||||||||||||
Mytiloides mytiloidiformis | X |
|
||||||||||||
Mytiloides puebloensis | X |
|
||||||||||||
Mytiloides ratonensis | X |
|
||||||||||||
Mytiloides scupini | X |
|
||||||||||||
Mytiloides striatoconcentricus | X |
|
||||||||||||
Mytilus sp. | X |
|
||||||||||||
Neithia irregularis | X |
|
||||||||||||
Neithia sp. | X |
|
||||||||||||
Nemodon eufaulensis | X |
|
||||||||||||
Nicaisolopha lugubris | X |
|
||||||||||||
Nucula sp. | X | X |
|
|||||||||||
Nuculidae indet. | X |
|
||||||||||||
Ostrea beloiti | X |
|
||||||||||||
Ostrea congesta | X | X |
|
|||||||||||
Ostrea contracta | X |
|
||||||||||||
Ostrea cretacea | X |
|
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Ostrea elegantula | X |
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Ostrea falcata | X |
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Ostrea glabra | X |
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Ostrea Johnsoni | X |
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Ostrea larva nasuta | X |
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Ostrea plumosa | X | X |
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Ostrea pratti | X | X |
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Ostrea satellites | X |
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Ostrea cf. O. veleniana | X |
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BIVALVIA (Continued) | ||||||||||||||
CLAMS, OYSTERS, MUSSELS | Ostrea subspatulata | X | X |
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Ostrea tecticostata | X |
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Paranomia sp. | X |
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Pecinidae indet. | X | X |
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Pecten cliffwoodensis | X |
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Pecten roemeri | X |
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Pecten subalpinus | X |
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Pecten texanus | X |
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Pedalion sp. | X |
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Pholadomya cf. P. coahuilensis | X |
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Pholadomya sanctisabae | X | X |
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Pholadomya sp. | X |
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Pholadomyidae indet. | X |
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Platyceramus americanus | X |
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Platyceramus cf. mantelli | X |
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Platyceramus platinus | X |
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Platyceramus sp. | X |
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Pleuromyidae indet. | X |
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Porocystis globularis | X |
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Protocardia multistriata | X |
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Protocardia texana | X | X |
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Protocardium sp. | X | X |
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Pseudoperna congesta | X |
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Pseudoperna n. sp. + | X |
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Pseudoperna sp. | X |
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Pteria sp. | X |
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Pycnodontae aucella | X |
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Pycnodontae sp. | X | X | X | X | X | X |
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Radiolites austinensis | X |
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Radiolites sp. | X |
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Radiolitidae indet. | X | X |
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Sphenoceramus digitatus | X |
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Sphenoceramus sp. | X |
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Spondylus cf. S. guadalupae | X |
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Spondylus sp. | X |
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Striarca poguei | X |
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Tapes chihuahuaensis | X | X |
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Tapes sp. | X |
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Tellenia elliptica | X |
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Tellenia simplex | X |
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Tellina sp. | X | X |
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Teredo irregularis | X |
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Teredo sp. | X |
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Teredolites sp. | X | X |
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Thracia gracilis | X |
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Thracia sp. | X |
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Trapezium truncatum | X |
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Trigonia bartrami | X |
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Trigonia sp. | X | X | X | X |
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Unio sp. | X | X | X |
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Venericardia sp. | X |
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Veneridae indet. | X |
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Veniella carolinensis | X |
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Veniella conradi | X |
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Veniella mullinensis | X |
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GASTROPODA | ||||||||||||||
SNAILS, WHELKS | Amauropsis sp. | X |
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Anchura kiowana | X |
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Anchura sp. | X |
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Aporrhais cf. A. subfusiformis | X |
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Aporrhais tarrantensis | X | X |
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Buccinopsis greenensis | X |
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Buccinopsis globosa | X |
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Buccinopsis parryi | X |
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Campeloma vetulum | X |
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Cerithidea indet. | X | *Cooper et al. current research | ||||||||||||
Cerithium sp. | X |
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certhiid gastropods | X |
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Cithara sp. | X |
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Gastropoda indet. (fresh water) | X | X | * |
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Gastropoda indet. ( |